Start with the animal the words actually name: a small creature that gets by in a place where almost nothing does. It lives in a hole, comes out when the ground cools, knows exactly where everything is, and needs very little. Every other use of desert rat takes that small tough survivor and hangs it on a person who has lived in the desert long enough to be like one.

先看字面上真的在講的那隻動物:一隻在幾乎沒東西活得下去的地方活下來的小生物。牠住在洞裡,等地面涼了才出來,對整片地熟得不得了,需要的東西非常少。desert rat 其他的用法,可以想成都是把這隻小小的、耐得住的生物,套在一個在沙漠裡待夠久的人身上。

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Second World War desert soldier · noun

A nickname for a soldier who served in the desert of North Africa in the Second World War, in the British 7th Armoured Division; the sources also count Australian soldiers who served with them.

沙漠之鼠(二戰北非英軍)